TRANSPORTATION OF LIVING MATERIALS -2,2

Capillarity is the action that causes water to rise in narrow tubes. Xylem vessels have a narrow lumen which makes it possible for water to rise in them by capillarity.
Capillarity is made possible by cohesion and adhesion forces. Cohesion is the attraction between like molecules. It makes the water molecules Stick to each other. Adhesion is attraction between different molecules. It causes water molecules to adhere to the xylem vessels.

Root pressure
Root   pressure   pushes   water   and   dissolved mineral salts upwards from the root. This happens because the cells of the endodermis push mineral It’s into the xylem. This increases osmotic pressure in the xylem thereby creating a force that moves the water and dissolved minerals up the xylem vessel. When a plant is cut, fluid oozes out the remaining stem (Stump). This is proof of root pressure in plant.


TRANSIPIRATION
Transpiration is the process by which plants lose water through the stomata in the leaves.
Water flows from the roots to the leaves through the xylem vessels. It enters the spongy mesophyll by osmosis. The spongy mesophyll has substomatal air spaces where water enters as water vapor. As a result, the concentration of water vapor in the substomatal air spaces becomes higher than the concentration of water vapor in the air. This causes water to diffuse into the atmosphere through the stomata.


         
                       Movement of water through leaves
Note: Another process known as guttation also occurs in plants. It is the process by which plants lose water as droplets through special glands found where veins are in contact with the leaf margin. Guttation is different from transpiration in that transpiration is the loss of water vapor mainly through the plant's stomata. Guttation occurs mostly at night or in plants growing in wet areas
Types of transpiration
There are three types of transpiration:
  1. Stomatal   transpiration   occurs   through the stomata on the leaves. It accounts for approximately 90% of the water lost by plants.
  2. Cuticular   transpiration   happens   through the cuticle of leaves. The cuticle is a waxy layer that covers the surface of leaves. A thick cuticle prevents excessive loss of water.
  3. Lenticular transpiration takes place through the lenticels. Lenticels are pores found on the bark of stems or roots in woody plant

Factors affecting the rate of transpiration
The rate of transpiration is affected by plant features as well as environmental factors.
Plant features
Plant features include the following:
(a)    The size of leaves; a large leaf has more stomata than a small leaf. Therefore, plants with large leaves lose more water than those with smaller leaves,
(b)   An extensive root system:Plants that have extensive roots absorb more water and can therefore lose more water than those with few roots.
(c)    Leaf cuticle: A thick cuticle resists water loss by transpiration while a thin cuticle makes water loss by transpiration easier.
(d)   Number of stomata: The more stomata a leaf have, the faster the rate of transpiration and vice versa.
(e)   Position of stomata:Stomata on the upper surface of the leaf lose water more easily than those on the lower surface. If a plant has leaves with more stomata on the upper surface, the rate of transpiration is faster than in        a plant that has Leaves with more stomata on the lower leaf surface.
(f)    Size of substomatal air spaces: Larger air spaces allow for a faster rate of transpiration because the leaves can hold more water vapor. Smaller substomatal air spaces slow down the rate of transpiration.
(g)   Sunken stomata: Sunken stomata occur in pits. They are not exposed to moving air so they slow down transpiration rate.
(h)   "Epidermal hairs: Epidermal hairs trap water on the surface of the leaves, thus preventing water

Environmental factors
(a)    Temperature: Transpiration rates go up as the temperature goes up. Higher temperatures cause the stomata to open and release water into [the atmosphere. Lower temperatures cause the stomata to close.
(b)   Relative humidity:As the relative humidity of the surrounding air rises, the transpiration rate falls. It is easier for water to evaporate into dry air than into air saturated with moisture.
(c)   Wind and air movement: Increased movement of the air around a plant results in a higher transpiration rate. As water transpires from a leaf, the water saturates the air surrounding the leaf. If there is no wind, the air              does not move, thus 11raising the humidity of the air around the leaf. Wind moves the air causing the more saturated air close to the leaf to be replaced by drier air.
(d)   Availability of soil moisture:When moisture is lacking in the soil, plants begin to senesce (age prematurely) resulting in leaf loss and reduced transpiration. Also, less water is absorbed by the roots when the soil is dry.
(e)   Light: Increased sunlight increases the rate of photosynthesis in the guard cells, causing them to become turgid and open the stomata. Higher light intensity also increases the plant's internal temperature and hence                    increases the rate of transpiration.
(f)    Atmospheric pressure: When atmospheric pressure is low, for example at high altitudes, plants lose water more easily. The rate of transpiration is reduced in areas with high atmospheric pressure.

Significance of transpiration
  1. It helps to maintain transpirational pull which is important for maintaining a constant stream of water between the roots and the leaves.
  2. Transpiration enables the loss of excess water from the plant,
  3. It helps to cool the plant and enables absorption and distribution of water and mineral salts.

Summary:
  1. The vascular system in plants is made up of xylem and phloem tissues.
  2. Xylem transports water and" mineral salts from the roots to all parts of the plant.
  3. Phloem transports manufactured food from the site of photosynthesis to all parts of the plant.
  4. The distribution of vascular bundles is different in roots and stems and in dicotyledonous and monocotyledonous plants.
  5. Root hairs are extensions of the epidermal cells of the root. They absorb water and mineral salts from the soil.
  6. Water is absorbed from the soil by osmosis.
  7. Mineral salts are absorbed from the soil by active transport.
  8. Water and dissolved minerals move up thexylem by transpiration pull, capillarity and root pressure.
  9. Transpiration is the process by which plants lose excess water through their leaves. Transpiration is important because it:

  • Helps   to maintain the transpirational stream.
  • Enables the loss of excess water.
  • Enables absorption and distribution of water and mineral salts in a plant.
  • Helps to cool the plant.

  1. Transpiration is affected by the features ofthe plant and environmental factors. The features of the plant include: leaf size, size of root system, size of leaf cuticle, size of air spaces, number and position of stomata and whether the stomata are sunken or not, and the presence of epidermal hairs.
  2. Environmental factors include the amounts of moisture in air, temperature, and air movement, availability of soil moisture, light and atmospheric pressure.





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